Our research group germinated in January 2005 and now includes high school interns, undergraduates, graduate students, technical support staff, and visiting researchers from around the world. If you'd like to learn more about volunteer positions, independent study opportunities, or graduate assistantships in our group (and/or in the Mycological Herbarium), please contact Betsy Arnold.

Principal Investigator

A. Elizabeth (Betsy) Arnold

Professor, School of Plant Sciences and Department of Ecology and Evolutionary Biology
Curator, Robert L. Gilbertson Mycological Herbarium, part of the University of Arizona Campus Herbarium

PhD, Ecology and Evolutionary Biology, University of Arizona (2002)
Ecology, evolution, and systematics of fungal endophytes; evolution of symbioses

CV: click here


Dr. Naupaka Zimmerman

Gordon and Betty Moore Postdoctoral Fellow, Life Sciences Research Foundation

PhD, Ecology, Stanford University (2013)

Biogeography, fungi-host interactions, and comparative genomics of foliar fungi

CV: click here
Dr. Joseph Spraker

PhD, Plant Pathology, University of Wisconsin, Madison (2016)

Biology of endohyphal bacteria and their impacts on fungal endophyte interactions with plants and secondary
metabolism; art-science interfacial learning.
Dr. Camilo Zalamea

Postdoctoral researcher, based at the University of Illinois and Smithsonian Tropical Research Institute, Panama
Tropical ecology

Graduate students

Elizabeth (Liz) Bowman

NSF Graduate Research Fellow and PhD student, Plant Pathology

Plant-fungal interactions as a function of climate change
Yu-Ling Huang

PhD student, Plant Sciences

Plant-fungal coevolution

CV: click here
Gavin Lehr

MS student, General Biology

Biology instructor, Sahuarita High School

Fungal communities associated with invasive plants
Shuzo Oita

PhD student, Plant Pathology

Fungal ecology
Justin Shaffer

PhD student, Plant Pathology (co-advised by David Baltrus)

Diversity and ecological effects of endohyphal bacteria associated with tropical seed-associated fungi; fungal ecology

CV:click here
Emma Holland

PhD student, Genetics

Phenotypic effects of endohyphal bacteria
Alison Harringon

PhD Student, Ecology and Evolutionary Biology
Aasiya Hamzazai

PhD Student, Plant Sciences
Technical assistants

Anyang Ndobegang

Fungal ecology and microbiology
Carolina Sarmiento

Based at the University of Illinois and Smithsonian Tropical Research Institute, Panama
Ming-Min Lee

BS, Biology, Oregon State University (2002)
MS, Entomology, University of Arizona (2008)
Kayla Garcia

BS, Microbiology, University of Arizona (2014)

Ellen Pat Photo-1.PNG
Ellen Pat
Nathaniel Yang
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Jasmin Pratt
Sequoia Fischer
Emma Jong
Jose Orozco
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Maithili Khandekar

Haley Williams

Halina Siewiora

Brianna Talbot

Mycological Herbarium
Joe Myers
High school students
2016: Emily Burke, Leah Simpson

2015: Tahlía Segura, Adalee Martínez, Wyatt Mendez, Leila Murrieta

Former graduate students (lab alumni): Dr. Michele Hoffman, Plant Pathology, PhD: graduated 5/2010 - US Homeland Security • Dr. Jana U'Ren, PhD, Plant Pathology: graduated 12/2011 - postdoctoral fellow, ABE, University of Arizona • Dr. Mariana del Olmo Ruiz, PhD, Plant Pathology: graduated 8/2012 - postdoctoral fellow, Institute for Ecology, UNAM (Mexico) • Dr. Ellen Martinson, PhD, Ecology and Evolutionary Biology: graduated 12/2012 - postdoctoral fellow, University of Rochester • Dr. Mary Jane Epps, PhD, Ecology and Evolutionary Biology: graduated 12/2012 - postdocoral fellow, North Carolina State University • Mr. Dustin Sandberg, MS, Plant Pathology: graduated 5/2013 - plant pathologist, USDA/APHIS • Ms. Kayla Arendt, MS, Plant Pathology: graduated 1/2015 - exploring opportunities • Ms. Chantelle Khambholja, MS, Plant Pathology: graduated 5/2015 - veterinary school
Ms. Sarah Araldi-Brondolo, MS, Plant Pathology, industry: graduated 8/2016.

Former postdocs (lab alumni):
Dr. Naupaka Zimmerman, Biogeography, fungi-host interactions, comparative genomics of foliar fungi • Dr. Nicholas Garber, sustainable agriculture
• Dr. Jana U'Ren, Dimensions of Biodiversity/endophytes of the boreal biome • Dr. Sarah Higginbotham, fungal secondary metabolites.

Interns, trainees, and visitors from other institutions: 2016, Gerald Schneider, University of Utah • 2016, Peter Tellez, Tulane University • 2016, Melissa Norris, UA • 2016, Daniel Hayden, Oklahoma State • 2015, Felix (Nattapol) Kraisitudomsook, Whitman College • 2014, Paulina Ramos, University of Sinaloa • 2014, Joao Paulo Toledo, Sao Paulo, Brazil • 2013, Erin Spear, University of Utah • 2013, Julian Gonzales III, Ramiro Garza, Kevin Thompson -- University of Texas, Pan-American • 2013, Carolina Sarmiento, Smithsonian Tropical Research Institute • 2013, Camilo Zalamea, Smithsonian Tropical Research Institute • 2012, Adriana Corrales, University of Illinois • 2012, Julien Ponchart, Universite Laval • 2012, Peter Cerda, University of Texas Pan American • 2012, Edelio Bazan, University of Texas Pan American • 2011, 2012, Brittany Peña, University of Texas Pan American • 2010-2011, Ochana Otto ('Otto'), Tucson High Magnet School • 2010-2013, Wes MacDonald, MacDonald Home School • 2010, Rowena Dolino, teacher at Tsehootsooi Middle School, Navajo Nation (AZ START program) • 2010, Jim Uomoto, teacher at Pueblo High Magnet School, Tucson (AZ START program) • 2010-2011, Frankie Orozco, student at Tucson High Magnet School, Tucson (Pima County) • 2010, Abby Cochrane, student at Catalina Foothills High Magnet School, Tucson (NSF) • 2010, Jessica Bolaños, SENACyT Panama • 2010, Seth Kauppinen, UC Berkeley • 2010, Sarah Higginbotham, Smithsonian Tropical Research Institute • 2010-2011, Sheri Steidl, Yavapai Community College (NSF) • 2009, Rowena Dolino, teacher at Tsehootsooi Middle School, Navajo Nation (AZ START program) • 2009, Catherina Caballero-George, SENACyT Panama • 2009, Magaly de Chial, Universidad de Panama • 2008- • 2011, Demetra Kandalepas, Louisana State University • 2008, Steve Uyeda, teacher at Sunnyside High School and instructor at Pima Community College (AZ START program) • 2008, Alex (JJ) Delgado, first-year student at Pima Community College (Arizona Biology Network) • 2006-2007, Cheyenne Weeks-Galindo, Pima Community College (tropical seed-associated fungi) (Arizona Biology Network) • 2006, Courtney Kluger, Duke University (tropical seed-associated fungi) (NSF REU) • 2005, 2007, Mary Shimabukuro, Dine College (fungal endophytes of ponderosa pine) (Arizona Biology Network)
Other lab alumni (and current professional status):
Ashton Leo • Michael J. Taylor, graduate school • Jose A. Bermudez • Carl Patterson-Markowitz • Sean Quigley • Thomas Gleason • Jorge Ramos, teaching molecular biotechnology in Mexico • Kayla Garcia • James deVore, research intern, Smithsonian Tropical Research Institute • Celine Bui • Wes MacDonald, UBRP student • Cole Steen, sustainable agriculture • Nick Massimo, in graduate school • Thao Truong, pharmacy school • Jakob Riddle, graduate school • Christie Moss, medical school • Brigid Heffernan, privately employed • Lorna Battista, privately employed • Nandi Devan, ecology and conservation biology/education • Brett Baxter, employed in medical field • Chan Jung, medical school • Jamie Moy, graduate school • Courtney Klopper, graduate school • Brittany Wohl, privately employed • Syed Ali Raza, medical school • Janka Vanova, pharmacy school • Emily Hendershot, privately employed • Patrick Campbell, pharmacy school • Barbara Beauchamp, teaching science in economically challenged schools • Barney Gilley, applying to graduate programs • Abby Cochrane, high school intern...now a UA undergraduate • Darren Stensrud, privately employed • Andrea Woodard, privately employed • Max Xiong, privately employed • Phornpoj Intarakamhang, applying to graduate programs • Erica Untch, teacher • Doug Mahana, graduate school • Jamal Alafifi, in graduate school • Yujin (Youchin) Huh, pharmacy school • Chan Maketon, graduate school • Dylan Grippi, graduate school • Lindsay Higgins, graduate school; now privately employed in scientific translation • Linh Huynh, pharmacy school • Ming-Min (Ming) Lee, graduate school • Elan Snitkin, privately employed • Rebecca Porter, privately employed • Julia Maddison, graduate school • Vanessa Chicharello, privately employed • Lindsay Cook, privately employed • Anita Bhakta, privately employed • Lauren Dominick, privately employed • Thaddeus Metz, privately employed • Adrian Ramirez, privately employed • Ethan Posey, UA student • Kiona Brown, privately employed • Jason Ong, privately employed • Ben Wilder, graduate school • Felicia Quintana, privately employed • Hilary Brown, privately employed • Melissa Kohmetscher, privately employed • Jean Tsai, graduate school • Patricia Espiritu, privately employed

Volunteers, Diné College project: 2006: Amanda Way, Liang Shen, Kelly Farrell (Duke University) and Claire Heinitz (UA). • 2007: Rebecca Porter (Duke University), Mariana del Olmo R., Fabiola Santos R., and Christine Coursodon (UA)...and thanks to Ming-Min Lee, Julia Maddison, Michele Hoffman, and Jana U'Ren for help with this project throughout its duration!
Students conducting lab rotations: Emma Holland (Genetics): endohyphal bacteria • Ryan Wallace (ABBS): endohyphal bacteria • Neill Prohaska (EEB): endophytes • Breonna Smith (ABBS): endolichenic fungi • Yu-Ling Huang (ABBS): effects of wildfire on fungal endophytes • Mana Ohkura (PLP): diversity of endophytes and endohyphal bacteria from the Navajo Nation • Simon Stump (EEB): Fungal communities associated with Erodium cicutarium, an invasive plant in SE, Arizona • Amritha Wickramage (PLP): diversity and specificity of fungi from the Chuska Mountains, Navajo Nation • David Jarvis (PLS/IGERT in Genomics): evolution of ecological modes in the Ascomycota • Jana U'Ren (PLS/IGERT in Genomics): evolutionary origins of tropical seed-associated fungi • Mary Jane Epps (EEB): diversity and ecological roles of nectar-inhabiting microbes • Mariana del Olmo Ruiz (PLP): estimating diversity of tropical foliar endophytes • Libby Landeen (EEB): endolichenic bacteria • Michele Hoffman (PLP): endophytes of Cupressaceae
Joe Myers, Trevor Mock, and Carl Patterson-Markowitz
Joe Myers, Trevor Mock, and Carl Patterson-Markowitz